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students presenting poster



Payne JL and Heim NA. 2020. Body size, sampling completeness, and extinction risk in the marine fossil record. Paleobiology 46:23-40.

Heim NA, *Bakshi S, *Buu L, *Chen S, *Heh S, *Jain A, *Noll C, *Patkar A, *Rizk N, *Sundararajan S, *Villante I, Knope ML, and Payne JL. 2020. Respiratory and circulatory anatomy constrain size evolution in solitary marine macrofauna. Paleobiology.  

*high school intern


McClain CR, Heim NA, Knope ML, and Payne JL. 2018. Is biodiversity energy-limited or unbounded? A test in fossil and modern bivalves. Paleobiology 44:385-401.


Heim, N., Payne, J., Finnegan, S., Knope, M., Kowalewski, M., Lyons, S., McShea, D., Novack-Gottshall, P., Smith, F., Wang, S.(2017). Hierarchical complexity and the size limits of life. Proceedings of the Royal Society B: Biological Sciences 284(1857): 20171039.


Payne, J., Bush, A., Chang, E., Heim, N., Knope, M., Pruss, S.(2016). Extinction intensity, selectivity and their combined macroevolutionary influence in the fossil record.Biology Letters 12(10): 20160202.

Payne, J., Bush, A., Heim, N., Knope, M., McCauley, D.(2016). Ecological selectivity of the emerging mass extinction in the oceans. Science 353(6305): 1284–1286.

Smith, F., Payne, J., Heim, N., Balk, M., Finnegan, S., Kowalewski, M., Lyons, S., McClain, C., McShea, D., Novack-Gottshall, P., Anich, P., Wang, S.(2016). Body size evolution across the Geozoic. Annual Reviews of Earth and Planetary Sciences 44(1): 523–553.

Keating-Bitonti, C., Payne, J.(2016). Physicochemical controls on biogeographic variation of benthic foraminiferal test size and shape. Paleobiology 42(4): 595–611.


Heim NA, Knope ML, Schaal EK, Wang SC, and Payne JL. 2014. Cope’s rule in the evolution of marine animals. Science 347:867-870. doi:10.1126/science.1260065

Zhang, Z., Augustin, M., Payne, J. (2015). Phanerozoic trends in brachiopod body size from synoptic dataPaleobiology 41(3): 491–501.


Harnik PG, Fitzgerald PC, Payne JL, and Carlson SJ. 2014. Phylogenetic signal in extinction selectivity in Devonian terebratulide brachiopods. Paleobiology 40:675–692. doi:10.1666/14006. doi:10.1666/14006


Caval-Holme F, Payne JL, and Skotheim JM. 2013. Constraints on the adult-offspring size relationship in protists. Evolution 67:3537–3544. doi:10.1111/evo.12210

Payne JL, Jost AB, Wang SC & Skotheim JM. 2013. A shift in the long-term mode of foraminiferan size evolution caused by the end-Permian mass extinction. Evolution 67:816-827. doi:10.1111/j.1558-5646.2012.01807.x.


Rego BL, Wang SC, Altiner D, & Payne JL. 2012. Within- and among-genus components of size evolution during mass extinction, recovery, and background intervals: a case study of Late Permian through Late Triassic foraminifera. Paleobiology 38:627–643. doi:10.1666/11040.1.

Payne JL, Groves JR, Jost AB, Nguyen T, Moffitt SE, Hill TM & Skotheim JM 2012. Late Paleozoic fusulinoidean gigantism driven by atmospheric hyperoxia. Evolution 66:2929–2939. doi:10.1111/j.1558-5646.2012.01626.x.

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Finnegan S, McClain CM, Kosnik MA & Payne JL. 2011. Escargots through time: an energetic comparison of marine gastropod assemblages before and after the Mesozoic Marine Revolution. Paleobiology 37:252–269. doi:10.1666/09066.1.

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Payne JL, Boyer AG, Brown JH, Finnegan S, Kowalewski M, Krause RA, Lyons SK, McClain CR, McShea DW, Novack-Gottshall PM, Smith FA, Stempien JA & Wang SC. 2009. Two-phase increase in the maximum size of life on Earth over 3.5 billion years reflects biological innovation and environmental opportunity. Proceedings of the National Academy of Sciences of the United States of America 106:24-27. doi:10.1073/pnas.0806314106.

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